Acacia is a genus of shrubs and trees belonging to the subfamily Mimosoideae of the family Fabaceae, first described in Africa by the Swedish botanist Carl Linnaeus in 1773. Many non-Australian species tend to be thorny, whereas the majority of Australian acacias are not. They are pod-bearing, with sap and leaves typically bearing large amounts of tannins and condensed tannins that historically in many species found use as pharmaceuticals and preservatives.
Acacias are also known as thorntrees, whistling thorns or wattles, including the yellow-fever acacia and umbrella acacias.
The genus Acacia previously contained roughly 1300 species, about 960 of them native to Australia, with the remainder spread around the tropical to warm-temperate regions of both hemispheres, including Europe, Africa, southern Asia, and the Americas. However, in 2005 the genus was divided into five separate genera. The name Acacia was retained for the majority of the Australian species and a few in tropical Asia, Madagascar and Pacific Islands. Most of the species outside Australia, and a small number of Australian species, were reclassified into Vachellia and Senegalia. The two final genera, Acaciella and Mariosousa, only contain about a dozen species from the Americas each.
Description
The leaves of acacias are compound pinnate in general. In some species, however, more especially in the Australian and Pacific islands species, the leaflets are suppressed, and the leaf-stalks (petioles) become vertically flattened, and serve the purpose of leaves. These are known as phyllodes. The vertical orientation of the phyllodes protects them from intense sunlight, as with their edges towards the sky and earth they do not intercept light so fully as horizontally placed leaves. A few species (such as Acacia glaucoptera) lack leaves or phyllodes altogether, but possess instead cladodes, modified leaf-like photosynthetic stems functioning as leaves.
The plants often bear spines, especially those species growing in arid regions. These sometimes represent branches which have become short, hard and pungent, or sometimes leaf-stipules. Acacia armata is the Kangaroo-thorn of Australia and Acacia erioloba is the Camelthorn of Africa.
Acacia seeds can be difficult to germinate. Research has found that immersing the seeds in various temperatures (usually around 80 °C) and manual seed coat chipping can improve yields to approximately 80 percent.
Similar mutualisms with ants occur on Acacia trees in Africa, such as the Whistling Thorn acacia. The acacias provide shelter for ants in the hollow stipules and nectar in extrafloral nectaries for their symbiotic ants such as Crematogaster mimosae. In turn, the ants protect the plant by attacking large mammalian herbivores and stem-boring beetles that damage the plant.
Uses and Symbolism
Acacia farnesiana is used in the perfume industry due to its strong fragrance. The use of Acacia as a fragrance dates back centuries.
The Acacia is used as a symbol in Freemasonry, to represent purity and endurance of the soul, and as funerary symbolism signifying resurrection and immortality. The tree gains its importance from the description of the burial of Hiram Abiff, the builder of King Solomon's Temple in Jerusalem.
Egyptian mythology has associated the acacia tree with characteristics of the tree of life (see the article on the Myth of Osiris and Isis).
Perhaps the most famous acacia is the Arbre du Ténéré in Niger. The reason for the tree's fame is that it used to be the most isolated tree in the world, approximately 400 km (249 mi) from any other tree. The tree was knocked down by a truck driver in 1973.
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